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Mary Katherine (Amanda Seyfried)
MK is a smart, spirited and headstrong 17-year-old who returns to her childhood home to connect with her estranged father, but has little patience for his endless stories of the ''tiny people'' who live in the woods. That all changes when she is magically transported into the Leafmen's world and must join the fight to save it in order to get home.
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Nod (Josh Hutcherson)
The handsome Nod is all about bucking the rules and flying solo, and there's no room for that kind of individualism in the Leaf Men, the warriors who fight to protect mother nature. But when MK enters the picture and the stakes are raised, Nod discovers what it takes to be a true hero.
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Bomba (Jason Sudeikis)
In his quest to study this civilization of tiny people he's never seen, this quirky, absent-minded professor has grown out of touch with his now teenage daughter, MK. His home in the middle of the forest is full of bizarre homemade gadgets, but Bomba soon comes to figure out what's really important to him.
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Ronin (Colin Farrell)
The battle-hardened leader of the Leafmen lives by the creed ''many leaves, one tree,'' and wouldn't think twice about putting his own life on the line for someone he cares about. When the forest and his beloved Queen come under siege, he's the one everyone else turns to to lead the way.
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Queen Tara (Beyoncé Knowles)
She is not only the leader of this magical world, but also the life force of the forest. She presides over it all with respect, compassion and humor, particularly when it comes to her close friend Ronin. When Tara finds herself in danger, her unique connection with nature gives her powerful allies.
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Grub (Chris O'Dowd) and Mub (Aziz Ansari)
This slug and snail duo may be low on the evolutionary food chain, but they have a pretty high sense of themselves. Mub, a self-characterized ''ladies' man,'' and Grub, a wannabe Leafman, are the caretakers of the royal pod that will bloom into the future Queen or King of the forest.
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He's a tough-guy toad who wields some power in the underbelly of the forest and knows how to play to the winning side.
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Nim Galuu (Steven Tyler)
At first glance, Nim is the consummate party animal. But this larger than life caterpillar a key behind-the-scenes player thanks to his wisdom and practical know-how.
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Mandrake (Christoph Waltz)
Gifted, or cursed, with the ability to bring destruction to anything he touches, Mandrake and his Boggan minions are the arch-nemeses of the Leafmen. Mandrake is tired of hiding in the shadows, and with his son by his side, he plans the ultimate revenge ? to claim the forest he believes should have always been his.